The lower shaft of a feather, specifically the region lacking barbs.

A pen made from a feather.

(by extension) Any pen.

A thin piece of bark, especially of cinnamon or cinchona, curled up into a tube.

The pen of a squid.

(music) The plectrum with which musicians strike the strings of certain instruments.

(music) The tube of a musical instrument.

(weaving) Something having the form of a quill, such as the fold or plain of a ruff, or a spindle, or spool, upon which the thread for the woof is wound in a shuttle.

To pierce with quills. (Usually in the passive voice, as be quilled or get quilled.)

(figurative) To write.

To form fabric into small, rounded folds.

To decorate with quillwork.

(US and Canada, especially Appalachia and the Prairies, transitive) To subject (a woman who is giving birth) to the practice of quilling (blowing pepper into her nose to induce or hasten labor).